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A perennial herb associated with a variety of wetland habitats including ponds, rivers, canals, marshes, fens, fen-meadows, wet woodland and dune-slacks. It also grows on coastal boulder beaches in Scotland. Generally lowland, but reaching 365 m at High Cup Gill (Westmorland).

Norfolk Skullcap

A perennial herb which was naturalised in a woodland near Brandon (W. Norfolk). Lowland.

Lesser Skullcap

A perennial herb of wet heaths, bogs, marshes and moist, heathy woodlands on acidic, oligotrophic or slightly mesotrophic soils. 0-440 m (Woodhead, Cheshire).